0.273 0.306 0.255 0.142 -0.184 0.395 0.373 0.135 -0.288 0.155 0.158 0.307 0.228 0.167 0.419 -0.170 0.153 0.089 0.150 0.095 0.290 0.240 0.222 0.187 -0.102 0.420 0.152 -0.298 Mean annual temperature Temperature variation Mean annual precipitation Precipitation variation Soil N Year Soil P Soil K N addition P addition K addition Temporal species turnover R 2 = 0.186 Leaf nitrogen R 2 = 0.525 Leaf phosphorus R 2 = 0.320 Leaf potassium R 2 = 0.550 SLA R 2 = 0.105